Carabid Beetles: Their Evolution, Natural History, and by George E. Ball (auth.), Terry L. Erwin, George E. Ball,

By George E. Ball (auth.), Terry L. Erwin, George E. Ball, Donald R. Whitehead, Anne L. Halpern (eds.)

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Mammals and amphibians). One group is often characterized as rapidly evolving and "speciating", tachytelic (Simpson, 1953), the other as evolving slowly (bradytelic). Such intergroup comparisons, if made for a limited time frame and between homogeneous groups, may demonstrate intrinsic differences between groups in their ability or opportunity to evolve. , see Whitehead, 1972]) are themselves valuable generalizations. They can be used in comparisons among different groups (as noted above) and also in gross testing for consistency between phylogenetic and zoogeographic reconstructions for the group; but they are usually too general to provide insights into the details of evolutionary mechanisms.

1974. Classification and comparative biology of the seed beetle genus Caryedes Hummel (Coleoptera: Bruchidae). Transactions of the American Entomological Society 100: 341-436. Lindroth, C. H. 1961-1969. The Ground-Beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Parts 1-6. Opuscala Entomologica, xlviii + 1192 pp. 1961, Part 2, Suppl. 20: 1-200; 1963, Part 3, Suppl. 24:201-408; 1966, Part 4, Suppl. 29:409-648; 1968, Part 5, Suppl. 33:649-944; 1969, Part 6, Suppl. 34:945-1192; 1969, Part I, Suppl.

My South American material was insufficient to test these ideas properly, but the general patterns of distribution and variation seem to fit very well. Among members of the genus, R. stigma (Linnaeus) is the most widespread, best represented, and most varied (Whitehead, 1976: fig. 142). Some of the populations form continuous character clines, some are geographically disjunctive, some are morphologically disjunctive from geographically proximate populations, and some are even sympatric and divergent in terms of character displacement.

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